Our heroic trio of Esau, Seneca, and Attar take the fight to the savage apes who have been besieging the remote, embattled township. They successfully follow them through the jungle back to their lair, but they're not prepared for what they find. The savage apes' home is, in fact, the lost ruins of the first Ape City abandoned centuries ago! While a hidden chapter from the history of the Planet of the Apes has been uncovered, another mystery is discovered: who or what were the Old Gods?